Puff, C. and Chamchumroon, V. (2014) “Thai Rubiaceae with hooks and thorns”, Thai Forest Bulletin (Botany), (31), pp. 65–74. available at: https://li01.tci-thaijo.org/index.php/ThaiForestBulletin/article/view/24405 (accessed: 2 April 2025).