Addition of Liopeltis frenatus (Günther, 1858) and Cyclophiops multicinctus (Roux, 1907) to the herpetofauna of Thailand (Squamata: Colubridae)
Keywords:
distribution, new snake records, northern Thailand, range extensions, Southeast AsiaAbstract
This contribution reports the first records for Thailand of two colubrid snake species, both from the eastern part of the country’s northern region. A living specimen of Liopeltis frenatus (Günther, 1858) was observed and photographed in submontane forest at an elevation of 1700 m in eastern Nan Province. A fresh and intact road-killed (DOR) specimen of Cyclophiops multicinctus (Roux, 1907) was collected in the low hills of eastern Uttaradit Province. The total range of both species is plotted in locality dot maps. The locality and habitat of the new records in Thailand are discussed in biogeographic perspective.
References
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Bain, R. H. and Hurley, M. M. 2011. A biogeographic synthesis of the amphibians and reptiles of Indochina. Bulletin of the American Museum of Natural History, no. 360: 1-138.
Boulenger, G. 1894. Catalogue of the Snakes in the British Museum (Natural History), Vol. 2. London, British Museum. 382 pp.
Bourret, R. 1935a. Notes Herpétologiques sur l’Indochine Française. X. Les Serpents de la station d’altitude du Tam-dao. Bulletin Général de l’Instruction Publique, Hanoi (No. 8, April 1935): 1-13. (in French)
Bourret, R. 1935b. Notes Herpétologiques sur l’Indochine Française. XI. Les Serpents de la station d’altitude du Tam-dao. Bulletin Général de l’Instruction Publique, Hanoi (No. 9, May 1935): 1-8. (in French)
Bourret, R. 1937a. Notes Herpétologiques sur l’Indochine Française. XIII. Serpents récemment récoltés au Tonkin et en Annam. Bulletin Général de l’Instruction Publique, Hanoi (No. 9, May 1937): 27-36. (in French)
Bourret, R. 1937b. Notes Herpétologiques sur l’Indochine Française. XV. Lézards et Serpents reçus au Laboratoire des Sciences Naturelles de l’Université au cours de l’année 1937. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Général de l’Instruction Publique, Hanoi (December 1937): 68-80. (in French)
Bourret, R. 1939a. Notes Herpétologiques sur l’Indochine Française. XVII. Reptiles et Batraciens reçus au Laboratoire des Sciences Naturelles de l’Université au cours de l’année 1938. Descriptions de trois espèces nouvelles. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Général de l’Instruction Publique, Hanoi (No 6, February 1939): 18-26. (in French)
Bourret, R. 1939b. Notes Herpétologiques sur l’Indochine Française. XVIII. Reptiles et Batraciens reçus au Laboratoire des Sciences Naturelles de l’Université au cours de l’année 1939. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Général de l’Instruction Publique, Hanoi (No. 4, December 1939): 5-39. (in French)
Bourret, R. 1939c. Notes Herpétologiques sur l’Indochine Française. XIX. La Faune Herpétologique des Stations d’altitude du Tonkin. Bulletin Général de l’Instruction Publique, Hanoi (No. 4, December 1939): 41-47. (in French)
Bourret, R. 1939d. Notes Herpétologiques sur l’Indochine Française. XX. Liste des Reptiles et Batraciens actuellement connus en Indochine Française. Bulletin Général de l’Instruction Publique, Hanoi (No. 4, December 1939): 49-60. (in French)
Campden-Main, S. M. 1970. A Field Guide to the Snakes of South Vietnam. Washington DC, Smithsonian Institute. 114 pp.
Chuaynkern, Y., Duangkae, P., Pongcharoen, C., Chuaynkern, C. and Horsin, L. 2014. Opisthotropis spenceri Smith, 1918 (Serpentes: Natricidae): the third and fourth specimens. Journal of Wildlife in Thailand 21(1): 1-14.
Cox, M. J., Hoover, M. F., Chanhome, L. and Thirakhupt, K. 2012. The snakes of Thailand. Bangkok, Chulalongkorn University Museum of Natural History. 845 pp.
Das, I. 2010. A field guide to the reptiles of Thailand and South-East Asia. Bangkok, Asia Books. 376 pp.
Das, I. 2012. A naturalist’s guide to the snakes of Thailand and Southeast Asia. Bangkok, Asia Books. 160 pp.
Das, I., Dattagupta, B. and N. C. Gayen. 1998. History and catalogue of reptile types in the collection of the Zoological Survey of India. Journal of South Asian Natural History 3(2): 121-172.
Deuve, J., 1970. Serpents du Laos. Paris, O. R. S. T. O. M., 251 pp.
Gardner, S., Sidisunthorn P. and Anusarnsunthorn, V. 2000. A Field Guide to the Forest Trees of Northern Thailand. Bangkok, Kobfai Publishing Project. 545 pp.
Geisler, P., Nguyen, T. Q., Poyarkov, N. A. and Böhme. W. 2011. New records of snakes from Cat Tien National Park, Dong Nai and Lam Dong provinces, southern Vietnam. Bonn zoological Bulletin 60(1): 9-16.
Günther, A., 1858. Catalogue of Colubrine snakes of the British Museum. London. 281 pp.
Hauser, S. 2017. On the validity of Pareas macularius Theobald, 1868 (Squamata: Pareidae) as a species distinct from Pareas margaritophorus (Jan in Bocourt, 1866). Tropical Natural History 17(1): 147-174.
Krishna, M., Parimal, C., Ray, C., Sarma, K. and Kumar, A. 2012. Notes on less known Günther’s Reed Snake Liopeltis frenatus (Günther, 1858). NeBIO 3(3): 126-127.
Luu, V. Q., Nguyen, T. Q., Pham, C. T., Dang, K. N., Vu, K. N., Miscovic, S., Bonkowski, M. and Ziegler, T. 2013. No end in sight? Further new records of amphibians and reptiles from Phong Nha – Ke Bang National Park, Quang Binh Province, Vietnam. Biodiversity Journal 4(2): 285-300.
Murphy, J. C., Chan-ard, T., Mekchai, S., Cota, M. and Voris, H. K. 2008. The rediscovery of Angel’s stream snake, Paratapinophis praemaxillaris Angel, 1929 (Reptilia: Serpentes: Natricidae). The Natural History Journal of Chulalongkorn University 8(2): 169-183.
Nguyen, V. S., Ho, T. C. and Nguyen, Q. T. 2009. Herpetofauna of Vietnam. Frankfurt am Main, Editions Chimaira. 761 pp.
Orlov, N. L., Murphy, R. W. and Papenfuss, T. J. 2000. List of snakes of Tam-Dao mountain ridge (Tonkin, Vietnam). Russian Journal of Herpetology 7(1): 69-80.
Orlov, N. L., Ryabov, S. A., Nguyen, V. S. and Nguyen, Q. T. 2003. New records of poorly known snakes of Vietnam. Russian Journal of Herpetology 10(3): 217-240.
Pauwels, O. S. G., David, P., Nutphand, W. and Chimsunchart, C. 2001. First record of Xenochrophis punctulatus (Günther, 1858) (Serpentes: Colubridae: Natricinae) from Thailand. Hamadryad 26(2): 247-252.
Pauwels, O. S. G., David, P., Chimsunchart, C., and Thirakhupt, K. 2003. Reptiles of Phetchaburi Province, Western Thailand: a list of species, with natural history notes, and a discussion on the biogeography at the Isthmus of Kra. The Natural History Journal of Chulalongkorn University 3(1): 23-53.
Pauwels, O. S. G., Kunya, K., David, P., Waengsothorn, S. and Sumontha, M. 2009a. On the occurrence of Amphiesma khasiense (Serpentes: Natricidae) in Thailand. Hamadryad 34(1): 75-79.
Pauwels, O. S. G., Kunya, K., David, P. and Sumontha, M. 2009b. First record of the Yunnan Keelback Sinonatrix yunnanensis Rao & Yang, 1998 (Serpentes: Natricidae) from Thailand. Salamandra 45 (3): 165-169.
Pawar, S. and Birand, A. (2001) A survey of amphibians, reptiles and birds in Northeast India. CERC Technical Report 6. Mysore, Centre for Ecological Research and Conservation. 118 pp.
Pham, A. V., Nguyen, S. L. H. and Nguyen, T. Q. 2014. New records of snakes (Squamata: Serpentes) from San La Province, Vietnam. Herpetology Notes 7: 771-777.
Roux, Jean, 1907. Diagnosen neuer Reptilien aus Asien und Amerika. Zoologischer Anzeiger, Leipzig 31: 762-65.
Santisuk, T., 1989. The monotypic family Bretschneideraceae newly recorded for Thailand. Natural History Bulletin of the Siam Society 37(2): 173-176.
Sharma, R. C., 2003. Handbook Indian Snakes. Kolkata, Zoological Survey of India. 292 pp.
Sharma, R. C., 2007. Fauna of India and the adjacent countries. Reptilia Volume-III (Serpentes). Kolkata, Zoological Survey of India. 410 pp.
Smith, M. A., 1940. The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma. Records of the Indian Museum 42(3): 465-486.
Smith, M. A. 1943. The fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. London, Taylor & Francis. 583 pp.
Stuart, B. L., Sok, K. and Neang, T. 2006. A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology 54: 129-155.
Teynié, A. and David, P. 2013. Amphibiens et reptiles des formations karstiques du Laos. Bulletin de la Société Herpétologique de France 148: 451-489. (in French)
Teynié, A. and Hauser, S. 2017. First record of Parafimbrios lao Teynié, David, Lottier, Le, Vidal et Nguyen, 2015 (Squamata: Caenophidia: Xenodermatidae) for Thailand. Russian Journal of Herpetology 24(1): 41-48.
Teynié, A., Lottier, A. and David, P. 2017. Azemiops feae Boulenger, 1888 (Squamata: Viperidae) et cinq autres additions notables pour la faune des serpents du Laos. Bulletin de la Société Herpétologique de France 162: 39-54. (in French)
Teynié, A., Nguyen, T. Q., Lorvelec, O., Piquet, A., Lottier, A. and David, P. 2014. Amphibiens et reptiles du Laos: nouvelles données nationales et provinciales. Bulletin de la Société Herpétologique de France 151: 21-52. (in French)
Tillack, F., Scheidt, U. and Ihle, T., 2006. First record of Blakeway’s mountain snake, Plagiopholis blakewayi Boulenger, 1893 from Thailand, with remarks on the distribution of Plagiopholis nuchalis Boulenger, 1893 (Reptilia: Squamata: Colubridae, Pseudoxenodontinae). Veröffentlichungen des Naturkundemuseums Erfurt, 25: 181-186.
Vasaruchapong, T., Laoungbua, P., Tangrattanapibul, K., Tawan, T. and Chanhome, L. 2017. Protobothrops mucrosquamatus (Cantor, 1839), a Highly Venomous Species Added to the Snake Fauna of Thailand (Squamata: Viperidae). Tropical Natural History 17(2): 111-115.
Vogel, G. 2009. On the distribution of Pareas hamptoni (Boulenger, 1905) in Thailand (Serpentes: Pareatinae). Russian Journal of Herpetolology 17(3): 219-222.
Vogel, G. and Hauser, S. 2013. Addition of Ptyas nigromarginata (Blyth, 1854) (Squamata: Colubridae) to the Snake Fauna of Thailand with Preliminary Remarks on Its Distribution. Asian Herpetological Research 4(3): 166-181.
Vogel, G., van Rooijen, J. and Hauser, S. 2012. A new species of Dendrelaphis Boulenger, 1890 (Squamata: Colubridae) from Thailand and Myanmar. Zootaxa 3392: 35-46.
Wall, F., 1908. Notes on a collection of snakes from the Khasi Hills, Assam. Journal of the Bombay Society 18: 312-327.
Wall, F. 1925. Notes on Snakes collected in Burma in 1924. Journal of the Bombay Natural History Society 30: 805-821.
Wallach, V., Williams, K. L. and Boundy, J. 2014. Snakes of the World. A Catalogue of Living and Extinct Species. Boca Raton (Florida), CRC Press. 1227 pp.
Xiao, Z., Li, Z.-C. and Qing, N., 2010. Discovery of Cyclophiops multicinctus in Guangdong Province. Chinese Journal of Zoology 2010(3).
Yang, D. and Rao, D., 2008. Amphibia and reptilia of Yunnan. Kunming, Yunnan Science and Technology Press. 499 pp. (in Chinese)
Ziegler, T., Hendrix, R., Thanh, V. N., Vogt, M., Foster, B. and Kien, D. N. 2007. The diversity of a snake community in a karst ecosystem in the central Truang Son, Vietnam, with an identification key. Zootaxa 1493: 1-40.
Ziegler, T., David, P., Mirallès, A., Doan, K. V. and Nguyen, T. Q. 2008. A new species of the snake genus Fimbrios from Phong Nha-Ke Bang National Park, Truong Son, central Vietnam (Squamata: Xenodermatidae). Zootaxa 1729: 37-48.
Ziegler, T., Tran, D. T. A., Nguyen, T. Q., Pwerl, R. G. B., Wirk, L., Kulisch, M., Lehmann, T., Rauhaus, A., Nguyen, T. T., Khac, Le, Q. K. and Vu, T. N. 2014. New amphibian and reptile records from Ha Giang Province, northern Vietnam. Herpetology Notes, 7: 185-201.
Bain, R. H. and Hurley, M. M. 2011. A biogeographic synthesis of the amphibians and reptiles of Indochina. Bulletin of the American Museum of Natural History, no. 360: 1-138.
Boulenger, G. 1894. Catalogue of the Snakes in the British Museum (Natural History), Vol. 2. London, British Museum. 382 pp.
Bourret, R. 1935a. Notes Herpétologiques sur l’Indochine Française. X. Les Serpents de la station d’altitude du Tam-dao. Bulletin Général de l’Instruction Publique, Hanoi (No. 8, April 1935): 1-13. (in French)
Bourret, R. 1935b. Notes Herpétologiques sur l’Indochine Française. XI. Les Serpents de la station d’altitude du Tam-dao. Bulletin Général de l’Instruction Publique, Hanoi (No. 9, May 1935): 1-8. (in French)
Bourret, R. 1937a. Notes Herpétologiques sur l’Indochine Française. XIII. Serpents récemment récoltés au Tonkin et en Annam. Bulletin Général de l’Instruction Publique, Hanoi (No. 9, May 1937): 27-36. (in French)
Bourret, R. 1937b. Notes Herpétologiques sur l’Indochine Française. XV. Lézards et Serpents reçus au Laboratoire des Sciences Naturelles de l’Université au cours de l’année 1937. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Général de l’Instruction Publique, Hanoi (December 1937): 68-80. (in French)
Bourret, R. 1939a. Notes Herpétologiques sur l’Indochine Française. XVII. Reptiles et Batraciens reçus au Laboratoire des Sciences Naturelles de l’Université au cours de l’année 1938. Descriptions de trois espèces nouvelles. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Général de l’Instruction Publique, Hanoi (No 6, February 1939): 18-26. (in French)
Bourret, R. 1939b. Notes Herpétologiques sur l’Indochine Française. XVIII. Reptiles et Batraciens reçus au Laboratoire des Sciences Naturelles de l’Université au cours de l’année 1939. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Général de l’Instruction Publique, Hanoi (No. 4, December 1939): 5-39. (in French)
Bourret, R. 1939c. Notes Herpétologiques sur l’Indochine Française. XIX. La Faune Herpétologique des Stations d’altitude du Tonkin. Bulletin Général de l’Instruction Publique, Hanoi (No. 4, December 1939): 41-47. (in French)
Bourret, R. 1939d. Notes Herpétologiques sur l’Indochine Française. XX. Liste des Reptiles et Batraciens actuellement connus en Indochine Française. Bulletin Général de l’Instruction Publique, Hanoi (No. 4, December 1939): 49-60. (in French)
Campden-Main, S. M. 1970. A Field Guide to the Snakes of South Vietnam. Washington DC, Smithsonian Institute. 114 pp.
Chuaynkern, Y., Duangkae, P., Pongcharoen, C., Chuaynkern, C. and Horsin, L. 2014. Opisthotropis spenceri Smith, 1918 (Serpentes: Natricidae): the third and fourth specimens. Journal of Wildlife in Thailand 21(1): 1-14.
Cox, M. J., Hoover, M. F., Chanhome, L. and Thirakhupt, K. 2012. The snakes of Thailand. Bangkok, Chulalongkorn University Museum of Natural History. 845 pp.
Das, I. 2010. A field guide to the reptiles of Thailand and South-East Asia. Bangkok, Asia Books. 376 pp.
Das, I. 2012. A naturalist’s guide to the snakes of Thailand and Southeast Asia. Bangkok, Asia Books. 160 pp.
Das, I., Dattagupta, B. and N. C. Gayen. 1998. History and catalogue of reptile types in the collection of the Zoological Survey of India. Journal of South Asian Natural History 3(2): 121-172.
Deuve, J., 1970. Serpents du Laos. Paris, O. R. S. T. O. M., 251 pp.
Gardner, S., Sidisunthorn P. and Anusarnsunthorn, V. 2000. A Field Guide to the Forest Trees of Northern Thailand. Bangkok, Kobfai Publishing Project. 545 pp.
Geisler, P., Nguyen, T. Q., Poyarkov, N. A. and Böhme. W. 2011. New records of snakes from Cat Tien National Park, Dong Nai and Lam Dong provinces, southern Vietnam. Bonn zoological Bulletin 60(1): 9-16.
Günther, A., 1858. Catalogue of Colubrine snakes of the British Museum. London. 281 pp.
Hauser, S. 2017. On the validity of Pareas macularius Theobald, 1868 (Squamata: Pareidae) as a species distinct from Pareas margaritophorus (Jan in Bocourt, 1866). Tropical Natural History 17(1): 147-174.
Krishna, M., Parimal, C., Ray, C., Sarma, K. and Kumar, A. 2012. Notes on less known Günther’s Reed Snake Liopeltis frenatus (Günther, 1858). NeBIO 3(3): 126-127.
Luu, V. Q., Nguyen, T. Q., Pham, C. T., Dang, K. N., Vu, K. N., Miscovic, S., Bonkowski, M. and Ziegler, T. 2013. No end in sight? Further new records of amphibians and reptiles from Phong Nha – Ke Bang National Park, Quang Binh Province, Vietnam. Biodiversity Journal 4(2): 285-300.
Murphy, J. C., Chan-ard, T., Mekchai, S., Cota, M. and Voris, H. K. 2008. The rediscovery of Angel’s stream snake, Paratapinophis praemaxillaris Angel, 1929 (Reptilia: Serpentes: Natricidae). The Natural History Journal of Chulalongkorn University 8(2): 169-183.
Nguyen, V. S., Ho, T. C. and Nguyen, Q. T. 2009. Herpetofauna of Vietnam. Frankfurt am Main, Editions Chimaira. 761 pp.
Orlov, N. L., Murphy, R. W. and Papenfuss, T. J. 2000. List of snakes of Tam-Dao mountain ridge (Tonkin, Vietnam). Russian Journal of Herpetology 7(1): 69-80.
Orlov, N. L., Ryabov, S. A., Nguyen, V. S. and Nguyen, Q. T. 2003. New records of poorly known snakes of Vietnam. Russian Journal of Herpetology 10(3): 217-240.
Pauwels, O. S. G., David, P., Nutphand, W. and Chimsunchart, C. 2001. First record of Xenochrophis punctulatus (Günther, 1858) (Serpentes: Colubridae: Natricinae) from Thailand. Hamadryad 26(2): 247-252.
Pauwels, O. S. G., David, P., Chimsunchart, C., and Thirakhupt, K. 2003. Reptiles of Phetchaburi Province, Western Thailand: a list of species, with natural history notes, and a discussion on the biogeography at the Isthmus of Kra. The Natural History Journal of Chulalongkorn University 3(1): 23-53.
Pauwels, O. S. G., Kunya, K., David, P., Waengsothorn, S. and Sumontha, M. 2009a. On the occurrence of Amphiesma khasiense (Serpentes: Natricidae) in Thailand. Hamadryad 34(1): 75-79.
Pauwels, O. S. G., Kunya, K., David, P. and Sumontha, M. 2009b. First record of the Yunnan Keelback Sinonatrix yunnanensis Rao & Yang, 1998 (Serpentes: Natricidae) from Thailand. Salamandra 45 (3): 165-169.
Pawar, S. and Birand, A. (2001) A survey of amphibians, reptiles and birds in Northeast India. CERC Technical Report 6. Mysore, Centre for Ecological Research and Conservation. 118 pp.
Pham, A. V., Nguyen, S. L. H. and Nguyen, T. Q. 2014. New records of snakes (Squamata: Serpentes) from San La Province, Vietnam. Herpetology Notes 7: 771-777.
Roux, Jean, 1907. Diagnosen neuer Reptilien aus Asien und Amerika. Zoologischer Anzeiger, Leipzig 31: 762-65.
Santisuk, T., 1989. The monotypic family Bretschneideraceae newly recorded for Thailand. Natural History Bulletin of the Siam Society 37(2): 173-176.
Sharma, R. C., 2003. Handbook Indian Snakes. Kolkata, Zoological Survey of India. 292 pp.
Sharma, R. C., 2007. Fauna of India and the adjacent countries. Reptilia Volume-III (Serpentes). Kolkata, Zoological Survey of India. 410 pp.
Smith, M. A., 1940. The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma. Records of the Indian Museum 42(3): 465-486.
Smith, M. A. 1943. The fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptilia and Amphibia. Vol. III, Serpentes. London, Taylor & Francis. 583 pp.
Stuart, B. L., Sok, K. and Neang, T. 2006. A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology 54: 129-155.
Teynié, A. and David, P. 2013. Amphibiens et reptiles des formations karstiques du Laos. Bulletin de la Société Herpétologique de France 148: 451-489. (in French)
Teynié, A. and Hauser, S. 2017. First record of Parafimbrios lao Teynié, David, Lottier, Le, Vidal et Nguyen, 2015 (Squamata: Caenophidia: Xenodermatidae) for Thailand. Russian Journal of Herpetology 24(1): 41-48.
Teynié, A., Lottier, A. and David, P. 2017. Azemiops feae Boulenger, 1888 (Squamata: Viperidae) et cinq autres additions notables pour la faune des serpents du Laos. Bulletin de la Société Herpétologique de France 162: 39-54. (in French)
Teynié, A., Nguyen, T. Q., Lorvelec, O., Piquet, A., Lottier, A. and David, P. 2014. Amphibiens et reptiles du Laos: nouvelles données nationales et provinciales. Bulletin de la Société Herpétologique de France 151: 21-52. (in French)
Tillack, F., Scheidt, U. and Ihle, T., 2006. First record of Blakeway’s mountain snake, Plagiopholis blakewayi Boulenger, 1893 from Thailand, with remarks on the distribution of Plagiopholis nuchalis Boulenger, 1893 (Reptilia: Squamata: Colubridae, Pseudoxenodontinae). Veröffentlichungen des Naturkundemuseums Erfurt, 25: 181-186.
Vasaruchapong, T., Laoungbua, P., Tangrattanapibul, K., Tawan, T. and Chanhome, L. 2017. Protobothrops mucrosquamatus (Cantor, 1839), a Highly Venomous Species Added to the Snake Fauna of Thailand (Squamata: Viperidae). Tropical Natural History 17(2): 111-115.
Vogel, G. 2009. On the distribution of Pareas hamptoni (Boulenger, 1905) in Thailand (Serpentes: Pareatinae). Russian Journal of Herpetolology 17(3): 219-222.
Vogel, G. and Hauser, S. 2013. Addition of Ptyas nigromarginata (Blyth, 1854) (Squamata: Colubridae) to the Snake Fauna of Thailand with Preliminary Remarks on Its Distribution. Asian Herpetological Research 4(3): 166-181.
Vogel, G., van Rooijen, J. and Hauser, S. 2012. A new species of Dendrelaphis Boulenger, 1890 (Squamata: Colubridae) from Thailand and Myanmar. Zootaxa 3392: 35-46.
Wall, F., 1908. Notes on a collection of snakes from the Khasi Hills, Assam. Journal of the Bombay Society 18: 312-327.
Wall, F. 1925. Notes on Snakes collected in Burma in 1924. Journal of the Bombay Natural History Society 30: 805-821.
Wallach, V., Williams, K. L. and Boundy, J. 2014. Snakes of the World. A Catalogue of Living and Extinct Species. Boca Raton (Florida), CRC Press. 1227 pp.
Xiao, Z., Li, Z.-C. and Qing, N., 2010. Discovery of Cyclophiops multicinctus in Guangdong Province. Chinese Journal of Zoology 2010(3).
Yang, D. and Rao, D., 2008. Amphibia and reptilia of Yunnan. Kunming, Yunnan Science and Technology Press. 499 pp. (in Chinese)
Ziegler, T., Hendrix, R., Thanh, V. N., Vogt, M., Foster, B. and Kien, D. N. 2007. The diversity of a snake community in a karst ecosystem in the central Truang Son, Vietnam, with an identification key. Zootaxa 1493: 1-40.
Ziegler, T., David, P., Mirallès, A., Doan, K. V. and Nguyen, T. Q. 2008. A new species of the snake genus Fimbrios from Phong Nha-Ke Bang National Park, Truong Son, central Vietnam (Squamata: Xenodermatidae). Zootaxa 1729: 37-48.
Ziegler, T., Tran, D. T. A., Nguyen, T. Q., Pwerl, R. G. B., Wirk, L., Kulisch, M., Lehmann, T., Rauhaus, A., Nguyen, T. T., Khac, Le, Q. K. and Vu, T. N. 2014. New amphibian and reptile records from Ha Giang Province, northern Vietnam. Herpetology Notes, 7: 185-201.
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2018-04-01
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HAUSER, S. 2018. Addition of Liopeltis frenatus (Günther, 1858) and Cyclophiops multicinctus (Roux, 1907) to the herpetofauna of Thailand (Squamata: Colubridae). Tropical Natural History. 18, 1 (Apr. 2018), 54–67.
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