Recent Discoveries in Zingiberaceae: A Review Highlighting New Species from Asia Over the Last Six Years
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Abstract
The increasing pace of habitat loss and environmental changes necessitates urgent research into the biodiversity of plant families such as Zingiberaceae. This review explores the discovery of new species within the Zingiberaceae family in Asia over the past six years, focusing on their distribution, characteristics, and the significance of these findings. The literature review involved searching various databases, including ScienceDirect, Scopus, Web of Science, Google Scholar, Google Books, and the Internet Archive, for references on new Zingiberaceae species. Specific search terms such as "new Zingiberaceae species," "discovery," "Asia," and "characterization" were used to identify pertinent articles. After collecting and analysing data, a total of 119 new species were identified across 11 countries, namely Cambodia, China, India, Indonesia, Laos, Malaysia, Myanmar, the Philippines, Taiwan, Thailand, and Vietnam. The genera include Alpinia, Boesenbegia, Curcuma, Etlingera, Globba, Hedychium, Kaempferia, Monolophus, Plagiostachys, Pleuranthodium, Sundamomum, Wurfbaiana, Conamomum, and Zingiber. Notably, the genus Curcuma was the most prolific, with 37 new species. Thailand emerged as the leading country in terms of new species discoveries. This comprehensive review highlights the diverse characteristics of these species and underscores the critical role of Zingiberaceae in ecological and economic contexts. The findings emphasize the importance of continued botanical exploration and research to enhance our understanding of biodiversity and to unlock potential applications in medicine, agriculture, and other industries.
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References
Ardi, W. H., & Ardiyani, M. (2016). Two new species of Alpinia (Zingiberaceae) from Sulawesi, Indonesia. Reinwardtia, 14(2), 311-316.
Ardiyani, M., Ardi, W. H., Hutabarat, P. W. K., & Poulsen, A. D. (2021a). Etlingera comosa, a new species (Zingiberaceae: alpinioideae) from Central Sulawesi. Reinwardtia, 20(2), 63-68.
Ardiyani, M., Ardi, W. H., Santoso, W., & Poulsen, A. D. (2021b). Etlingera tjiasmantoi (Zingiberaceae), a new species from Central Sulawesi. Reinwardtia, 19(2), 103-108.
Atmaja, M. B., Andila, P. S., Oktavia, G. A. E., & Merriansyah, H. (2023). Indonesian threatened Zingiberaceae: Exploring their potential traditional and modern uses. IOP Conference Series: Earth and Environmental Science, 1255, Article 012036. https://doi.org/10.1088/1755-1315/1255/1/012036
BPS. (2024). Produksi tanaman biofarmaka menurut provinsi dan jenis tanaman, 2024. https://www.bps.go.id/id/statistics-table/3/VVZNelkycEdWM2t5V2poTFltOVVURWR 0WWs1Mlp6MDkjMw==/produksi-tanaman-biofarmaka-menurut-provinsi-dan-jenis-tanaman--2023.html?year=2024
Bahishti, A. A. (2021). The importance of review articles & its prospects in scholarly literature. Extensive Reviews, 1(1), 1-6. https://doi.org/10.21467/exr.1.1.4293
Bai, L., Leong-Škorničková, J., & Xia, N. H. (2015). Taxonomic studies on Zingiber (Zingiberaceae) in China I: Zingiber kerrii and the synonymy of Z. menghaiense and Z. stipitatum. Gardens’ Bulletin Singapore, 67(1), 129-142. https://doi.org/10.3850/s2382581215000149
Banaticla-Hilario, M. C. N., & Altamirano, M. R. B. (2023). Conservation of IUCN threatened Zingiberaceae species in Tropical Asia: challenges, gaps and opportunities. In S. Ramamoorthy, I. E. Buot Jr., & C. Rajasekaran (Eds.). Plant diversity in biocultural landscapes (pp. 673-724). Springer Nature Singapore. https://doi.org/10.1007/978-981-19-8649-9_30
Bhaumik, M., Langhu, T., Dey, S., Deb, C. R., & Jamir, N. S. (2017). Two new species of Monolophus (Zingiberaceae) from India. Kew Bulletin, 72, 1-7. https://doi.org/10.1007/s12225-017-9687-4
Boonma, T. (2023). Curcuma suraponii sp. nov. (Zingiberaceae), a new species of Curcuma subgen. Curcuma from Thailand. Biodiversitas, 24(9), 4885-4895. https://doi.org/10.13057/biodiv/d240931
Boonma, T., Saensouk, S., & Saensouk, P. (2020). Two new species of Kaempferia L. (Zingiberaceae) from Thailand. Taiwania, 65(3), 371-381. https://doi.org/10.6165/tai.2020.65.371
Boonma, T., Saensouk, S., & Saensouk, P. (2021). Kaempferia nigrifolia (Zingiberaceae), a new species from Central Thailand. Rheedea, 31(1), 11-17.
Boonma, T., Saensouk, S., & Saensouk, P. (2022). Kaempferia sipraiana (Zingiberaceae), a new species from Thailand and a new record of Kaempferia pseudoparviflora for Myanmar. Biodiversitas Journal of Biological Diversity, 23(4), 2203-2211. https://doi.org/10.13057/biodiv/d230456
Chen, J., Tong, Y. H., & Xia, N. H. (2019). Curcuma yingdeensis (Zingiberaceae), a new species from China. Phytotaxa, 388(1), 98-106. https://doi.org/10.11646/phytotaxa.388.1.4
Chen, J., Ye, Y.-S., & Xia, N.-H. (2021). Curcuma ruiliensis (Zingiberaceae), a new species from Yunnan, China. Nordic Journal of Botany, 39(2), Article e02910. https://doi.org/10.1111/njb.02910
de Boer, H., Newman, M., Poulsen, A. D., Jane Droop, A., Fér, T., Hiền, L. T. T., Hlavatá, K., Lamxay, V., Richardson, J. E., Steffen, K., & Leong-Škorničková, J. (2018). Convergent morphology in Alpinieae (Zingiberaceae): Recircumscribing Amomum as a monophyletic genus. Taxon, 67(1), 6-36. https://doi.org/10.12705/671.2
de Castro, C. E. F., Gonçalves, C., Loges, V., Tavares, A. R., de Barros, F., & de Castro, A. C. R. (2023). Zingiber: synonymy, accepted and excluded species. Ornamental Horticulture, 29(2), 286-298. https://doi.org/10.1590/2447-536X.v29i2.2558
Debnath, S., Patil, S., & Vijayan, D. (2024). Boesenbegia ashihoi (Zingiberaceae), a new species from northeast India. Nordic Journal of Botany, 2024(5), Article e04319. https://doi.org/10.1111/NJB.04319
Deiji, N., Ahmad, B. N., Puranjoy, M., & Yogendra, K. (2020). Zingiber flavofusiforme (Zingiberaceae), a new record for the flora of India. Journal of Japanese Botany, 95(2), 102-105.
Dhillon, P. (2022). How to write a good scientific review article. FEBS Journal, 289(13), 3592-3602. https://doi.org/10.1111/febs.16565
Ding, H. B., Gong, Y. X., & Tan, Y. H. (2022). Globba depingiana (Zingiberaceae), a new species from Yunnan, China. Annales Botanici Fennici, 59(1), 57-60. https://doi.org/10.5735/085.059.0110
Ding, H. B., Ma, Y., & Tan, Y. H. (2023). Zingiber longii (Zingiberaceae), a new species from Southeast Yunnan, China. Taiwania, 68(3), 298-302. https://doi.org/10.6165/tai.2023.68.298
Ding, H. B., Quan, D. L., Zeng, X. D., Li, J. W., & Tan, Y. H. (2021). Zingiber calcicola (Zingiberaceae), a new species from a limestone area in south Yunnan, China. Phytotaxa, 525(1), 65-69. https://doi.org/10.11646/phytotaxa.525.1.8
Ding, H. B., Yang, B., Lu, X. Q., & Tan, Y. H. (2020a). Zingiber porphyrochilum (Zingiberaceae), a new species from Yunnan, China. Annales Botanici Fennici, 57(4-6), 197-201. https://doi.org/10.5735/085.057.0401
Ding, H. B., Yang, B., Maw, M. B., Win, P. P., & Tan, Y. H. (2020b). A new species and two new combinations of Monolophus (Zingiberaceae) from Indo-Burma. PhytoKeys, 138, 155-162. https://doi.org/10.3897/phytokeys.138.39217
Docot, R. V. A. (2020). Plagiostachys lourdesiae (Zingiberaceae), a new species from Mindanao, Philippines. Nordic Journal of Botany, 38(7), Article 02806. https://doi.org/10.1111/njb.02806
Docot, R. V. A., Domingo, C. B. M., Moran, C. B., Camangeg, L. M., & Poulsen, A. D. (2022). Wurfbainia rubrofasciata (Zingiberaceae), a new species from Palawan, Philippines. Webbia, 77(2), 277-283. https://doi.org/10.36253/jopt-13461
Docot, R. V. A., Gutierrez, K. D., Mamalias, R. E. E., Espino, N. B. R., Java, A. A. B., Dineros, C. D., & Mijares, E. M. L. (2019). Two new Zingiber species (Zingiberaceae) from Sorsogon, Philippines. Gardens’ Bulletin Singapore, 71(2), 459-475. https://doi.org/10.26492/gbs71(2).2019-14
Docot, R. V. A., Santiago, L. C. P., Funakoshi, H., & Lam, N. F. (2020). Two new species of Boesenbegia (Zingiberaceae) from Palawan, Philippines. Edinburgh Journal of Botany, 77(3), 377-390. https://doi.org/10.1017/S0960428620000098
Furmuly, A. M., & Azemi, N. (2020). A review on golden species of Zingiberaceae family: genus Curcuma. Science Proceeding Series, 2(2), 319-326.
Handayani, D., Sari, W. D. P., Purnama, D., Ritonga, Y. E., & Prakasa, H. (2022). Exploration of Zingiberaceae family in Tangkahan Ecotourism Forest, North Sumatra Province. AIP Conference Proceedings, 2659(1), Article 060016. https://doi.org/10.1063/5.0121872
Hu, X., Lu, J., Lao, Q., Wu, F., Li, Y, & Zhang, H. (2024). Inter-group variation of floral scent composition in two Hedychium groups (Zingiberaceae). https://ssrn.com/abstract=4951557
Huang, Z., Xie, L., Wang, H., Zhong, J., Li, Y., Liu, J., Ou, Z., Liang, X., Li, Y., Huang, H., Lin, Z., Zhang, K., Zhang, L., & Zheng, X. (2019). Geographic distribution and impacts of climate change on the suitable habitats of Zingiber species in China. Industrial Crops and Products, 138, Article 111429. https://doi.org/10.1016/j.indcrop.2019.05.078
Insisiengmay, O., Newman, M. F., & Haevermans, T. (2020). Two new species of Kaempferia L. (zingiberaceae) from cambodia and LAO PDR. European Journal of Taxonomy, 712, 1-15. https://doi.org/10.5852/ejt.2020.712
Jayakrishnan, T., Joe, A., Hareesh, V. S., & Sabu, M. (2021). Two new Zingiber (Zingiberaceae) species from Arunachal Pradesh, northeastern india. Taiwania, 66(1), 101-112. https://doi.org/10.6165/tai.2021.66.101
Jenjittikul, T., & Larsen, K. (2020). Two new species of Kaempferia (Zingiberaceae) from Thailand. The Routledge Handbook of English for Academic Purposes, 64, 403-415. https://doi.org/10.6165/tai.2020.65.371
Jenjittikul, T., & Ruchisansakun, S. (2020). Kaempferia albiflora (Zingiberaceae), a new species from Thailand. Kew Bulletin, 75(1), Article 13. https://doi.org/10.1007/s12225-020-9868-4
Jia, X. C., Li, J., Lu, G. H., & Wang, Y. Q. (2015). The concrete evidence of flexistyly in Plagiostachys: Pollination biology of a wild ginger on Hainan Island, China. Ecology and Evolution, 5(22), 5364-5371. https://doi.org/10.1002/ece3.1807
Juwitaningsih, T., Jahro, I. S., & Sari, S. A. (2020). Evaluation of north sumatera cardamom seed (Amomum compactum) extract as antibacterial and anticancer. Journal of Physics: Conference Series, 1485, Article 012019. https://doi.org/10.1088/17426596/1485/1/012019
Kaewsri, W., & Sangvirotjanapat, S. (2022). Six new species of Wurfbainia (Zingiberaceae) from Thailand. Edinburgh Journal of Botany, 79, Article 369. https://doi.org/10.24823/ejb.2022.369
Kajornjit, P., Saensouk, S., & Saensouk, P. (2018). Pollen morphology and leaf anatomy of genus Globba in Thailand. ScienceAsia, 44(3), 146-161. https://doi.org/10.2306/scienceasia1513-1874.2018.44.146
Karunarathne, P., Yakandawala, D., & Samaraweera, P. (2021). Fruit morphology helps identifying evolutionary groups in Alpinieae (Zingiberaceae): Inferences from phylogenetic analysis of gingers in Sri Lanka. Journal of the National Science Foundation of Sri Lanka, 49(3), 337-350. https://doi.org/10.4038/jnsfsr.v49i3.9820
Kiat, T. S., Pippen, R., Yusof, R., Rahman, N. A., IbrHIM, H., & Khalid, N. (2006). Screening of selected Zingiberaceae extracts for dengue-2 virus protease inhibitory activities. Sunway Academic Journal, 3, 1-7.
Kress, W. J., & Erickson, D. L. (2012). DNA barcodes : methods and protocols. Methods in Molecular Biology, 858, 3-8. https://doi.org/10.1007/978-1-61779-591-6_1
Kress, W. J., Prince, L. M., & Williams, K. J. (2002). The phylogeny and a new classification of the gingers (Zingiberaceae): evidence from molecular data. American Journal of Botany, 89(10), 1682-1696. https://doi.org/10.3732/ajb.89.10.1682
Lam, N. F., Ibrahim, H., Sam, Y. Y., Zakaria, R. M., & Poulsen, A. D. (2022). Two new species of Boesenbegia (Zingiberaceae), from Sabah, Malaysia. PhytoKeys, 211, 81-92. https://doi.org/10.3897/phytokeys.211.83985
Lee, J.-A., Lee, M.-Y., Seo, C.-S., Jung, D. Y., Lee, N.-H., Kim, J.-H., Ha, H., & Shin, H. K. (2010). Anti-asthmatic effects of an Amomum compactum extract on an ovalbumin (OVA)-induced murine asthma model. Bioscience, Biotechnology, and Biochemistry, 74(9), 1814-1818. https://doi.org/10.1271/bbb.100177
Lê, T.-H., Trịnh, T.-H., Ðỗ, N.-D., Nguyễn, V.-H., & Lý, N.-S. (2019). Zingiber vuquangense (Sect. Cryptanthium: Zingiberaceae): a new species from North central coast region, Vietnam. Phytotaxa, 388(4), 295-300. https://doi.org/10.11646/phytotaxa.388.4.5
Leong-Škorničková, J., Soonthornkalump, S., Niwesrat, S., & Lim, S. Q. (2022). Curcuma lindstromii (Zingiberaceae: Zingiberoideae), a new species from southeastern Thailand. Gardens’ Bulletin Singapore, 74(2), 243-250. https://doi.org/10.26492/gbs74(2).2022-09
Leong-škorničková, J., Soonthornkalump, S., & Thongbai, W. (2020a). Four new Curcuma species (Zingiberaceae) from Thailand.
Blumea: Biodiversity, Evolution and Biogeography of Plants, 65(3), 244-253. https://doi.org/10.3767/blumea.2021.65.03.09
Leong-Škornicková, J., Soonthornkalump, S., & Suksathan, P. (2020b). Curcuma cinnabarina and C. eburnea (Zingiberaceae: Zingiberoideae), two new species from Thailand. Edinburgh Journal of Botany, 77(3), 391-402. https://doi.org/10.1017/S0960428620000049
Li, R., Shine, L., Li, W., & Zhou, S. S. (2020). A new species of Zingiber (Zingiberaceae) from Natma Taung National Park, Chin State, Myanmar. PhytoKeys, 138, 131-137. https://doi.org/10.3897/phytokeys.138.46719
Lofthus, Ø., Newman, M. F., Jimbo, T., & Poulsen, A. D. (2020). The Pleuranthodium (Zingiberaceae) of Mount Wilhelm, Papua New Guinea. Blumea: Biodiversity, Evolution and Biogeography of Plants, 65(2), 95-101. https://doi.org/10.3767/blumea.2020.65.02.01
Lofthus, Ø. (2014). The genus Pleuranthodium (K. Schum.) R.M.Sm. (Zingiberaceae): taxonomy and phylogeny. AGRIS - International System for Agricultural Science and Technology.
Lý, N. S., Đỗ, Đ. G., Cao, N. G., Trương, B. V., Nguyễn, V. T., & Leong-škorničková, J. (2021). Zingiber magang and Z. tamii (zingiberaceae), two new species from central vietnam. Taiwania, 66(2), 232-240. https://doi.org/10.6165/tai.2021.66.232
Lý, N. S., Hoang, T. S., Insisiengmay, O., Haevermans, T., & Newman, M. F. (2022). Conamomum vietnamense (Zingiberaceae), a new species from Tay Nguyen, Vietnam. Phytotaxa, 531(2), 129-135. https://doi.org/10.11646/phytotaxa.531.2.5
Ma, X. Da, Wang, W. G., Gong, Q. B., Xu, G. H., Shi, J. P., & Shen, J. Y. (2021). Globba ruiliensis, a new species of zingiberaceae from Yunnan, China. Taiwania, 66(1), 31-34. https://doi.org/10.6165/tai.2021.66.31
Maknoi, C., Saensouk, S., Saensouk, P., Rakarcha, S., & Thammarong, W. (2021). Two new species of Curcuma L. (zingiberaceae) from Thailand. Biodiversitas, 22(9), 3910-3921. https://doi.org/10.13057/biodiv/d220937
Mazo, K. R. F. (2022). Two new species of Plagiostachys (Zingiberaceae) from Zamboanga Peninsula, Philippines. Taiwania, 67(2), 186-194. https://doi.org/10.6165/tai.2022.67.186
Meechonkit, P., & Picheansoonthon, C. (2021). Two new species of Kaempferia L. (Zingiberaceae) from Northern Thailand. Pakistan Journal of Botany, 53(6), 2221-2227. https://doi.org/10.30848/PJB2021-6(18)
Mohamad, S., Kalu, M., & Poulsen, A. D. (2020). A new species and a new combination of Sundamomum (Zingiberaceae) from Sarawak, Borneo. Kew Bulletin, 75(4), 3-8. https://doi.org/10.1007/s12225-020-09919-y
Mood, J.D., Veldkamp, J.F., Dey, S. & Prince, L.M. (2014). Nomenclatural changes in Zingiberaceae: Caulokaempferia is a superfluous name for Monolophus and Jirawongsea is reduced to Boesenbergia. Gardens' Bulletin Singapore 66(2), 215-231.
Nguyen, H. T., Nguyen, N. A., Averyanov, L., Nguyen, D. D., & Le, C. T. (2023a). Curcuma tuanii (Zingiberaceae) a new species of subgenus Ecomata from northern Vietnam based on morphological and molecular evidence. Acta Botanica Brasilica, 37(06), 1-13. https://doi.org/10.1590/1677-941X-ABB-2023-0028
Nguyen, D. D., Saensouk, S., Nguyen, van C., Nguyen, van K., Pham, T. T. D., Nguyen, D. K., & Nguyen, T. L. T. (2023b). Taxonomic notes of Alpinia subsect. Catimbium (Zingiberaceae) in Vietnam: The first record of Alpinia nobilis and description of a new species Alpinia hoangviet. Biodiversitas, 24(10), 5293-5301. https://doi.org/10.13057/biodiv/d241010
Nopporncharoenkul, N., & Jenjittikul, T. (2024). Taxonomic revision of some taxa in Kaempferia subgenus Protanthium (Zingiberaceae) revealing a new species from Thailand and two new synonyms. Blumea - Biodiversity, Evolution and Biogeography of Plants, 69, 16-26. https://doi.org/10.3767/blumea.2024.69.01.03
Nopporncharoenkul, N., Laongsri, W., & Jenjittikul, T. (2020). Two new species of Kaempferia subgenus Protanthium (Zingiberaceae) from northern Thailand. Nordic Journal of Botany, 38(2), Article e02633. https://doi.org/10.1111/njb.02633
Nopporncharoenkul, N., Somnoo, T., Tanming, W., & Maknoi, C. (2021). Kaempferia jenjittikuliae (Kaempferia subg. Protanthium: Zingiberaceae), a new, endangered species endemic to Thailand. Edinburgh Journal of Botany, 78, 1-13. https://doi.org/10.24823/ejb.2021.350
Odyuo, N., Roy, D., & Mao, A. (2019a). Zingiber dimapurense (Zingiberace), a new species from Nagaland, India. NeBIO, 10(2), 59-65.
Odyuo, N., Roy, D. K., Lyngwa, C., & Mao, A. A. (2019b). Zingiber perenense, a new species in Zingiber section Cryptanthium (Zingiberaceae) from Nagaland, India. Srinagarind Medical Journal, 34(2), 1865-1871.
Prince, L. M., Mood, J. D., Mandákova, T., & de Boer, H. J. (2024). Molecular phylogeny of Boesenbegia (Zingiberaceae) reveals shared characters and infrageneric division. Blumea - Biodiversity, Evolution and Biogeography of Plants, 69(1), 1-12. https://doi.org/10.3767/blumea.2024.69.01.01
Ragsasilp, A., Saensouk, P., & Saensouk, S. (2022). Ginger family from Bueng Kan Province, Thailand: Diversity, conservation status, and traditional uses. Biodiversitas, 23(5), 2739-2752. https://doi.org/10.13057/biodiv/d230556
Rakarcha, S., Saensouk, S., Maknoi, C., Wongnak, M., Thammarong, W., & Saensouk, P. (2022). Curcuma lampangensis and C. sabhasrii, two new species of the family Zingiberaceae from northern Thailand. Biodiversitas, 23(9), 4448-4459. https://doi.org/10.13057/biodiv/d230910
Range, C., Tad-o, R. V. B., Alafag, J. I., & Napaldet, J. T. (2022). Boesenbegia igorota (Zingiberaceae), a new species from the Cordillera Central Range, Northern Philippines. Taiwania, 3(67), 311-317. https://doi.org/10.6165/tai.2022.67.311
Rayhannisa, R., Zumaidar, Z., Amalia, A., Sari, W., & Saudah, S. (2024). Morphological characteristics of kecombrang (Etlingera elatior (Jack) R. M. Smith) in several regions in Aceh Province, Sumatra. Al-Kauniyah: Jurnal Biologi, 17(2), 450-459. https://doi.org/10.15408/kauniyah.v17i2.34900
Ruchisansakun, S., & Jenjittikul, T. (2023). Curcuma ignea (Zingiberaceae), a spectacular new species from Thailand. Edinburgh Journal of Botany, 80, 1-8. https://doi.org/10.24823/ejb.2023.1959
Sabu, M., & Hareesh, V. S. (2020). Hedychium mechukanum (Zingiberaceae), a new species from the eastern Himalayas, India. Gardens’ Bulletin Singapore, 72(2), 291-297. https://doi.org/10.26492/gbs72(2).2020-12
Saensouk, P., & Saensouk, S. (2021). Diversity, traditional uses and conservation status of zingiberaceae in Udorn Thani Province, Thailand. Biodiversitas, 22(8), 3083-3097. https://doi.org/10.13057/biodiv/d220801
Saensouk, S., & Saensouk, P. (2019). Kaempferia mahasarakhamensis, a new species from Thailand. Taiwania, 64(1), 39-42. https://doi.org/10.6165/tai.2019.64.39
Saensouk, S., Saensouk, P., Pasorn, P., & Chantaranothai, P. (2016). Diversity and uses of Zingiberaceae in Nam Nao National Park, Chaiyaphum and Phetchabun provinces, Thailand, with a new record for Thailand. Agriculture and Natural Resources, 50(6), 445-453. https://doi.org/10.1016/j.anres.2016.08.002
Saensouk, P., Boonma, T., & Saensouk, S. (2021a). Curcuma siamensis (Zingiberaceae, zingibereae), a new species of Curcuma subgen. ecomatae from southeastern thailand. Biodiversitas, 22(12), 5239-5246. https://doi.org/10.13057/biodiv/d221201
Saensouk, S., Boonma, T., & Saensouk, P. (2021b). A new species and a new record of Curcuma subgen. Curcuma (zingiberaceae) from northern thailand. Biodiversitas, 22(9), 3661-3670. https://doi.org/10.13057/biodiv/d220903
Saensouk, S., Boonma, T., & Saensouk, P. (2021c). View of six new species and a new record of Curcuma L. (Zingiberaceae) from Thailand. Biodiversitas, 22(4), 1658-1685. https://doi.org/10.13057/biodiv/d220410
Saensouk, S., Boonma, T., Thomudtha, A., Thomudtha, P., & Saensouk, P. (2021d). Short communication: Curcuma wanenlueanga (Zingiberaceae), a new species of subgenus Curcuma from Thailand. Biodiversitas, 22(7), 2988-2994. https://doi.org/10.13057/biodiv/d220752
Saensouk, P., Boonma, T., & Saensouk, S. (2022a). Curcuma nakhonphanomensis (Zingiberaceae), a new species from the lower Mekong River basin, northeastern Thailand. Biodiversitas, 23(11), 6040-6048. https://doi.org/10.13057/biodiv/d231159
Saensouk, P., Boonma, T., & Saensouk, S. (2022b). Curcuma pulcherrima (Zingiberaceae), a new rare species of Curcuma subgen. Ecomata from eastern Thailand. Biodiversitas, 23(12), 6635-6644. https://doi.org/10.13057/biodiv/d231262
Saensouk, P., Boonma, T., Rakarcha, S., Maknoi, C., Wongnak, M., & Saensouk, S. (2022c). Two new species of Curcuma subgenus Ecomata (Zingiberaceae: Zingibereae), from Central and Southwestern Thailand. Biodiversitas, 23(9), 4578-4588. https://doi.org/10.13057/biodiv/d230925
Saensouk, P., Saensouk, S., & Boonma, T. (2022d). Two new species of Kaempferia subgenus Kaempferia (Zingiberaceae: Zingibereae) from Thailand. Biodiversitas, 23(8), 4343-4354. https://doi.org/10.13057/biodiv/d230860
Saensouk, S., Boonma, T., & Saensouk, P. (2022e). Curcuma achrae (Zingiberaceae), a new species from Central Thailand. Rheedea, 32(1), 30-45. https://doi.org/10.22244/RHEEDEA.2021.32.01.03
Saensouk, P., Boonma, T., Maknoi, C., & Saensouk, S. (2023). Curcuma ubonensis (Zingiberaceae), a new species of Curcuma subgen. Hitcheniopsis from Eastern Thailand. Notulae Botanicae Horti Agrobotanici Cluj-Napoca, 51(4), 1-16. https://doi.org/10.15835/NBHA51413374
Saensouk, P., Saensouk, S., Boonma, T., & Rakarcha, S. (2024a). Kaempferia sakolchaii sp. nov. and K. phuphanensis var. viridans var. nov. (Zingiberaceae), two new taxa from Northeastern Thailand. Holticulturae, 10(5), 430. https://doi.org/10.3390/horticulturae10050430
Saensouk, P., Saensouk, S., Maknoi, C., & Boonma, T. (2024b). Curcuma borealis sp. nov. and C. retrocalcaria sp. nov.(Zingiberaceae): Two novel taxa from Northern Thailand. Horticulturae, 10(8), 787. https://doi.org/10.3390/horticulturae10080787
Saensouk, P., Saensouk, S., Boonma, T., Oo, W. P., Htet, N. M., Maknoi, C., Bongcheewin, B., Htway, N. N., & Minn, H. M. (2025). Two new records of Boesenbergia Kuntze (Zingiberaceae: Zingibereae) for the Flora of Myanmar. Biodiversitas, 26(1), 480-489. https://doi.org/10.13057/biodiv/d260147
Saha, K., Sinha, R. K., & Sinha, S. (2020). Distribution, cytology, genetic diversity and molecular phylogeny of selected species of Zingiberaceae–A review. Feddes Repertorium, 131(1), 58-68.
Sangvirotjanapat, S., Đăng, T. H., & Newman, M. (2020). Ten new species of Globba section Globba from continental South-East Asia. Thai Forest Bulletin (Botany), 48(2), 212-233. https://doi.org/10.20531/tfb.2020.48.2.15
Sangvirojtanapat, S., & Newman, M. (2023). Flora of Thailand: Zingiberaceae. In Flora of Thailand Vol. 16 (pp. 333-747). Forest Herbarium, Royal Forest Department.
Sedek, A. S., Mohamad, S., Abd-Aziz, N., Sarjuni, M. N. H., Hadzuha, N. H., & Hakim, A. M. A. (2023). Diversity of wild gingers (Zingiberaceae) in Southern Peninsular Malaysia: panti forest reserve and labis forest reserve. Pertanika Journal of Tropical Agricultural Science, 47(1), 1-14. https://doi.org/10.47836/pjtas.47.1.02
Selvaraj, D., Sarma, R. K., & Sathishkumar, R. (2008). Phylogenetic analysis of chloroplast matK gene from Zingiberaceae for plant DNA barcoding. Bioinformation, 3(1), 24-27. https://doi.org/10.6026/97320630003024
Seo, C. S., Lee, J. A., Jung, D., Lee, H. Y., Lee, J. K., Ha, H., Lee, M. Y., & Shin, H. K. (2011). Simultaneous determination of liquiritin, hesperidin, and glycyrrhizin by HPLC-photodiode array detection and the anti-inflammatory effect of Pyungwi-san. Archives of Pharmacal Research, 34(2), 203-210. https://doi.org/10.1007/s12272-011-0204-2
Sirirugsa, P. (1992). Taxonomy of the genus Kaempferia (Zingiberaceae) in Thailand. Thai Forest Bulletin, 19, 1-15.
Sirirugsa, P. (1999). Thai Zingiberaceae : species diversity and their uses. Pure and Applied Chemistry, 70(11), 23-27.
Sofian, F. F., Pambayun, G. W., Runadi, D., Susilawati, Y., Tjitraresmi, A., Herdiana, Y., & Astuti, E. P. (2019). Larvicidal activity of ethanol extract and essential oil from Zingiber aromaticum Val. rhizome against Aedes aegypti larvae. Journal of Pharmaceutical Sciences and Research, 11(1), 11-14.
Soonthornkalump, S., Kongphapa, J., Vianmana, S., Kunlapa, N., & Leong-Škorničková, J. (2022). Curcuma stahlianthoides (Zingiberaceae), a new species from northeastern Thailand dispersed by ants. Blumea: Journal of Plant Taxonomy and Plant Geography, 67(1), 71-75. https://doi.org/10.3767/blumea.2022.67.01.09
Soonthornkalump, S., Ongsakul, A., Dolaji, A., & Leongškorničková, J. (2020). Curcuma papilionacea (Zingiberaceae), an unusual new species from southern Thailand. Phytotaxa, 432(1), 11-16. https://doi.org/10.11646/phytotaxa.432.1.9
Souvannakhoummane, K., & Maknoi, C. (2014). Curcuma peramoena Souvann. and Maknoi (Zingiberaceae): a new species from Lao PDR. Thai Journal of Botany, 6(2), 125-130.
Souvannakhoummane, K., Lanorsavanh, S., & Sangvirotjanapat, S. (2023). Globba amicitia (Zingiberaceae: Globbeae), a new species from Phou Khao Khouay National Protected Area, Bolikhamxay Province, Laos. Thai Forest Bulletin Botani, 51(2), 157-163.
Subehan, Usia, T., Iwata, H., Kadota, S., & Tezuka, Y. (2006). Mechanism-based inhibition of CYP3A4 and CYP2D6 by Indonesian medicinal plants. Journal of Ethnopharmacology, 105(3), 449-455. https://doi.org/10.1016/j.jep.2005.12.001
Sungkawati, M., Hidayati, L., Daryono, B. S., & Purnomo. (2019). Phenetic analysis of Curcuma spp. in Yogyakarta, Indonesia based on morphological and anatomical characters. Biodiversitas, 20(8), 2340-2347. https://doi.org/10.13057/biodiv/d200832
Suryani, Y. (2022). Produksi energi pada mikroorganisme secara anaerob. Fisiologi Mikroorganisme. Gunung Djati Publishing.
Syahid, S. F., & Heryanto, R. (2017). Short communication: Morpho-agronomic characteristics of twelve accessions of white turmeric (Curcuma zedoaria) germplasm. Biodiversitas, 18(1), 269-274. https://doi.org/10.13057/biodiv/d180135.
Tanaka, N., & Aung, M. M. (2020). Taxonomic studies on Myanmar Zingiberaceae III: two new species of Zingiber (Sect. Cryptanthium) from Kayah State. Bulletin of the National Museum of Nature and Science. Series B, Botany, 46(1), 39-46.
Tanaka, N., Hoang, V., Tran, V., Kieu, T., Tram, T., Khanh, N., Tagane, S., Funakoshi, H., & Souladeth, P. (2023). A new species of Alpinia (Zingiberaceae: subgenus Alpinia subsect. Catimbium) from Laos and Vietnam. Bulletin of the National Science Museum Series B, 49(1), 25-32. https://doi.org/10.50826/bnmnsbot.49.1
Tanaka, N., Tagane, S., & Souladeth, P. (2020). Zingiber collinsii Mood & Theilade (Zingiberaceae), a newly recorded ginger from Laos. Thai Journal of Botany, 12(2), 105-111.
Trimanto, Dwiyanti, D., & Indriyani, S. (2018). Morfologi, anatomi dan uji histokimia rimpang Curcuma aeruginosa Roxb; Curcuma longa L. dan Curcuma heyneana Valeton dan Zipj. [Morphology, anatomical and histochemical rhizome of Curcuma aeruginosa Roxb; Curcuma longa L. and Curcuma heyneana Valeton and Zipj]. Berita Biologi, 17(2), 123-133.
Truong, L. H., Dang, T. H., Dat, N. Q., Trung, N. T. Q., & Gioi, T. (2019). Conamomum odorum, a new species of Zingiberaceae from central Vietnam. Academia Journal of Biology, 41(3), 55-59. https://doi.org/10.15625/2615-9023/v41n3.13671
Ujilestari, T., Martien, R., Ariyadi, B., Dono, N. D., & Zuprizal. (2018). Self-nanoemulsifying drug delivery system (SNEDDS) of Amomum compactum essential oil: Design, formulation, and characterization. Journal of Applied Pharmaceutical Science, 8(6), 14-21. https://doi.org/10.7324/JAPS.2018.8603
Wang, C.-M., Lin, Y.-C., & Tseng, Y.-H. (2020). Zingiber chengii (Zingiberaceae), a new species from Taiwan. PhytoKeys, 139, 1-11. https://doi.org/10.3897/phytokeys.139.37294
Watson, L., & Dallwitz, M. J. (1992). The families of flowering plants: descriptions, illustrations, identification, and information retrieval. https://www.worldcat.org/title/ families-of-flowering-plants-descriptions-illustrations-identification-and-information-retrieval/oclc/34189020
Yeh, C.-L., Chung, S.-W., Kuo, Y.-W., Hsu, T.-C., Leou, C.-S., Hong, S.-J., & Yeh, C.-R. (2012). A new species of Zingiber (Zingiberaceae) from Taiwan, China, based on morphological and molecular data. Journal of Systematics and Evolution, 50(2), 163-169. https://doi.org/10.1111/j.1759-6831.2011.00179.x
Yusmaniar, Wardiyah, Suprapti, T., & Junaedi. (2015). Antibacterial activity of the essensial oils of lempuyang wangi (Zingiber aromaticum Val.), lempuyang gajah (Zingiber zerumbet Sm), and lempuyang emprit (Zingiber amaricans Bl.) on three gram negative bacteria. Asian Journal of Applied Sciences, 3(2), 290-293.
Zhang, L.-X., Ding, H.-B., Li, H.-T., Zhang, Z.-L., & Tan, Y.-H. (2019). Curcuma tongii, a new species of Curcuma subgen. Ecomatae (zingiberaceae) from southern Yunnan, China. Phytotaxa, 395(3), 241-247. https://doi.org/10.11646/phytotaxa.395.3.9
Zode, R., Tagade, W., & Meshram, M. (2021). Zingiber capitatum Roxb - a new report for Gondia district, (Ms) India. In P. P. Umale, & D. K. Koche (Eds.). Current Updates in Life Science (pp. 302-307). Pandit Jawaharlal Nehru Study Center.